Lechuguilla Restoration: Techniques Learned in the Southwest Focus on Microbial Preservation

article and photos by Val Hildreth-Werker & Jim C. Werker

Pre-printed from NSS News April 1997

From February through November of 1996, cavers gave hundreds of hours to restoration efforts in Lechuguilla. Six restoration expeditions, led by Jim Werker and Val Hildreth-Werker, accomplished tasks in the Southwestern Branch. Special attention was concentrated on preserving pristine areas of the cave and on developing restoration techniques to minimize disturbance of biota.

Sadly, a decade of exploration has created well-worn paths of impact through the main trails of this world-class cave. Stray footprints have taken big bites out of the pristine beauty once harbored by Lechuguilla Cave. Some damage is permanent. Some areas can be repaired and restored to near-original state. Restoration is happening, one tiny chunk of misplaced mud at a time.

Lechuguilla's hard-earned wisdom is leading cavers to figure out better ways to cave softly ... and leave no trace. Cavers have always been first in finding better ways to protect cave resources. In the spirit of that perpetual cycle to do better, Lechuguilla is presenting us another gift ... improvements in conservation-minded caving techniques.

The cave has received a lot of obvious impact. Restoration seems an overwhelming task. Lechuguilla has more than 89 miles of surveyed passage abundant with breeches of pristine areas. The obvious question jumps forth. Where do we start? Perhaps by concentrating restoration efforts on one small section, then moving to the next small area ... the results will eventually add up to some degree of accomplishment. We approached the Cave Resource Office about restoration tasks in the Southwestern Branch of Lechuguilla.

MICROBIAL STUDIES IN LECHUGUILLA

Because microbial studies in Lechuguilla are yielding positive and important results, it is time we take precautions to decrease our human impact on the biota. We clearly need to figure out methods for minimizing impact on microbial communities while accomplishing restoration. Interest in the microbes of Lechuguilla was originally sparked by Kiym Cunningham's early investigations of folia and corrosion residue in which he noted the presence of fungal and bacterial structures. Through the study of microbial communities in sub-surface environments on earth, such as Lechuguilla, Penny Boston and NASA scientists are working to develop models for what life might be like on Mars.

The weight of the responsibility to approach restoration with respect for the microbiota is magnified by the fact that strongly positive results are coming from Larry Mallory's investigation of bacterial agents in Lechuguilla. He is identifying cave bacteria which produce chemical substances that are effective in fighting various forms of cancer. The impact of humans on the microbial community has been investigated by Diana Northup and Kathy Lavoie. In separate studies, they have each tested heavily traveled routes, camps, bivy sites, and urine dumps throughout the cave. To help cavers understand the importance of microbial preserves in caves, Northup, Lavoie, and Mallory have collaborated in composing a draft pamphlet describing ways we can avoid impacting microbial communities. Suggested guidelines in the draft are designed for Lechuguilla and may not be applicable to other types of caves.


Microbes in Caves

This is a draft version. Comments to Diana Northup will be appreciated.

Diana E. Northup1, Kathy Lavoie2, Larry Mallory3

MICROBES FOUND IN CAVES

In caves, we have microbes that are resident and microbes that are transient. Transient microbes ride into caves on air currents, in water flows, on insects, on bats, and through humans. Anything or anyone entering a cave may carry transient microbes. Resident microbes, on the other hand, occur as native inhabitants of a cave. Native microbes depend solely on the resources within the cave for survival, but transient microbes tend to thrive where abundant organic materials are available.

Microbes that live by using organic carbon are called chemoheterotrophs. Guano deposits, flowing water, and decaying wood provide organic rich matter for chemoheterotrophic microbes such as fungi. Most fungi found in caves fit this picture and are most likely transients. Bacteria can also be brought in caves as transients. However, current research shows that most native microbes in caves are bacteria.

Within a typical cave, any place with sufficient moisture might contain microbes. If there is a stream running into the cave, you might expect to find algae washed in from the surface, protozoa in the sediments at the bottom of the stream, and bacteria clinging to the surface of the water and air (bacteria like to hang out at interfaces where rock meets water and water meets air). The banks along a stream, with their deposits of sediment, would be home to fungal spores. These spores, the fungal equivalent of seeds, only grow when organic matter is encountered. Bacteria are abundant in deposits of soil or sediment, but most bacteria are dormant unless suitable food is present. Fungal spores and bacteria are found in the surface water film of limestone rock and calcite speleothems. These microbes may contribute to the formation and degradation of speleothems.

Two of the few places where you are likely to see colonies made up of billions and billions of bacterial cells is on limestone or lava tube surfaces. Those reflective white dots, clustered together in moist areas on cave ceilings and walls are actinomycete bacteria. Actinomycetes are responsible for the distinctive odor that caves and soils have, unlike musty basements whose odor is due to fungi. The air we breathe (you won't like this part!) contains millions of fungal spores and floating bacteria. Deep in caves, bacteria are found living in pools and dripwater. These bacteria are specialized to grow in very low nutrient, or oligotrophic, environments.

WHY MICROORGANISMS IN CAVES ARE IMPORTANT

Bacteria and fungi that make their home in caves are important for several reasons. Because of their long isolation from the surface and because of their existence in very low nutrient environments (we're not talking about the bat guano microbes here!), some cave microbes appear to have evolved to produce specialized chemical compounds, or toxins, with which to fend off neighboring microbes (their own version of assault rifles). These microbial chemical compounds may be useful to humans in the fight against disease or pollution. Preliminary results from Mallory's studies suggest that microbes demonstrating this sort of beneficial activity were collected from pristine sites that were rarely visited by humans.

Our knowledge of the microbial world in general is really quite limited and our knowledge of cave microbial diversity is even more limited. Thus, the potential exists to find novel microbes in caves. Investigation of such organisms may provide new details about the evolutionary relationships of bacteria and fungi.

The study of microbes in caves is also important in elucidating how speleothems are formed. There is good, although limited, evidence that microbes are involved in the formation of iron and manganese oxides, sulfur compounds, saltpeter deposits, and even calcium carbonate.

Finally, scientists are finding bacteria deep within rocks of the earth and in association with deep-sea hydrothermal vents. The deep subsurface environment is difficult to sample; thus, caves provide a more accessible avenue for studying these microbes.

HUMANS IMPACT MICROBES IN CAVES

The impact of humans on microbes in caves takes two forms. We import foreign (surface) microbes as we explore caves. Secondly, we bring additional organic matter into the cave in the form of skin, hair, food, lint, urine, and possibly even feces. You are shedding tens of thousands of skin fragments per minute! This additional organic matter does the most harm. Native cave microbes often live in very low nutrient environments and may not even be able to survive in richer environments. If we add too much organic matter, the cave habitat will cease to be a good place for native bacteria to live; and will become, instead, a good place for transient, surface microbes to thrive!

CONSERVING CAVE MICROBES

To decrease the number of foreign microbes you bring into the cave:

To limit the amount of human-associated organic matter entering the cave

(particularly in caves without streams):

In conserving microorganisms and the habitat in which they exist, balance is needed. Exploration teams often inform scientists of habitat that might contain microbes of interest. If the care taken by explorers of Lechuguilla Cave can serve as a model for other exploration endeavors, we can preserve microbes that may tell us much about the world in which we live. In order to preserve the more easily impacted microbes, we may wish to establish a few microbial preserves in remote areas of pristine caves and in areas with unique habitats.

If you have comments on the draft pamphlet, please contact Diana Northup at the Biology Department, University of New Mexico, Albuquerque, NM 87131.

1 Department of Biology, University of New Mexico, Albuquerque, NM 87131

2 Department of Biology, University of Michigan-Flint, Flint, MI 48502-2186

3 Biomes, 170 North Valley Road, Pelham, MA 01002-9766


AVOID IMPACTING MICROBIAL COMMUNITIES

Lechuguilla poses a different set of challenges for cave microorganisms. Unique microbes that require low-nutrient environments are sheltered by Lechuguilla. Because it is a deep cave harboring isolated pools and passages, specialized microbial communities inside Lechuguilla survive in the absence of surface nutrients. Many other caves receive major inputs of surface nutrients from streams and rivers. Unlike Lechuguilla, food is simply more plentiful where streams flow into caves and bring an abundance of carbon-loaded, nutrient-rich debris. In caves like Lechuguilla, where the carbon load is very low, microorganisms are adapted to living in low-nutrient environments. When cavers introduce skin, hair, food, mud, and other carbon-rich materials from the surface, the amount of nutrients in the cave environment may change.

With all the organic stuff we bring into caves, it is easy to understand that human presence might build up the nutrient base. Surface microbes have a better chance of surviving in caves when we introduce carbon-rich nutrients. When the food supply changes, the microbial community is likely to change. The surface organisms, enabled by the introduced food supply, may take over the environment. Native low-nutrient organisms may not be able to survive. Adding to or changing the nutrient base may favor the survival of surface microbes over the survival of native microbes. Microbial communities requiring low-nutrient environments could easily be destroyed by human impact.

Cavers doing restoration now have to be acutely aware of contamination issues! How do we work in areas and minimize our human impact? Preserving the microbiota is the major concern for restoration projects in Lechuguilla. How do we adapt common restoration practices for use in a giant microbial preserve? Following are descriptions of techniques used in Lechuguilla. More questions, answers, and better ideas will arise, but for 1996, here are examples of how we approached restoration with respect to the amazing diversity of sub-surface microbes.

RESTORATION WITH RESPECT TO MICROBIAL COMMUNITIES

First, we are using latex gloves for nearly every restoration task, especially in pristine or little traveled areas. Powder-free latex, vinyl, or rubber gloves are worn. Some people may be allergic to latex, though teams have not experienced ill-effects. Watch for rashes on the hands. Occasional severe reactions have been reported in medical literature. Holes and tears can be a problem. Some cavers rarely rip their gloves while others go through several pairs a day. Surgical gloves are reasonably comfortable, work well, fit right, and last longer than other choices.

Many restoration projects require water. In Lechuguilla, water is not carried into the cave for restoration tasks. It is carefully and conservatively taken from designated pools. Striving to take care in not cross-contaminating pools, water is used only in the area near the pool from which it came. Turkey basters are used to put pool water into spray bottles. Bottles, sprayers, basters, sponges, buckets, brushes, etc. are new or disinfected for each area of the cave. Because scientists have found differing microbial communities in pools only a few feet apart, we try to always identify and avoid possibilities for contamination.

To conserve water, simple filters are put to work. Restoration water and debris collected in sponges is squeezed into buckets and allowed time for the larger particles to settle. If the water does not clear, it is strained through a simple sponge filter. Foam or sponge material is compressed into a clean water bottle with the bottom cut off. Debris is caught around the edges of the sponge. Treated foam products are avoided since we do not know what chemicals they may add to the cave. Restoration water is filtered through this makeshift funnel and recycled for another dose of restoration work in the same area.

Mud sometimes collects on the bottoms of pools near trails due to foot travel. Hand pumps are used to collect the silt while water is filtered directly back into the pool. The makeshift funnel described above is held at the water outflow, allowing debris to be caught in the filter. For some pools, a rubber hose is attached to the pump. Two cavers can operate the device, one pumping and filtering, one directing the hose. The pump is disinfected before using it in a different pool.

We are investigating methods to disinfect equipment between uses. The following information has been provided by James Cokendolpher. Currently, chlorine bleach and hydrogen peroxide look like the disinfectants of choice. Both need a contact time of about 10 minutes; you can not just wipe them on and off and expect to kill much. Hydrogen peroxide rapidly breaks down into water and oxygen. There are two types of chlorine bleach: one is calcium hypochlorite and the other is sodium hypochlorite. They break down into sodium or calcium chloride (sodium chloride is table salt) and water. Both are highly corrosive and special care should be taken to keep them away from caving gear. Because of the toxicity of these compounds to microbes, they should be carried and stored in the cave in small containers. In that way, any spills will not wipe-out large numbers of desirable microbes. One tablespoon of bleach per gallon of water will kill many microbes! More research is needed to determine what residue is left by various disinfecting agents, what percentage is effective, what organisms are addressed, how long the agents continue to work, and how the cave is effected.

FLAGGING TRAILS IN LECHUGUILLA

Trail maintenance is an important aspect of restoration. When Lechuguilla trails were originally designated, flagging tape was laid in one foot lengths and a dotted line of trail was created. Often, only one side of trail was marked. Problems persisted with lost flagging, hard to find pathways, and too many misplaced footsteps. Based on years of observation, Carlsbad Caverns National Park made the decision that double flagged trails have positive and beneficial influence on the preservation of cave resources. Trails are now being double flagged throughout Lechuguilla. Double flagging--continuous lines of orange surveyors tape defining both sides of the trail--helps guide footsteps to stay on impacted areas. Double flagged trails may be visually more obtrusive. However, for the National Park Service, protection of the cave comes first, and experience has defined that double trails do help to conserve Lechuguilla's fragile, irreplaceable resources.

Laying flagging tape for trails takes a bit of skill development. There tends to be more to the process than expected. It is good to define techniques before entering the cave, and then work in small groups, each with an experienced flagging person directing the process.

Flagging is always "tied down" to prevent inadvertent movement of the tape. Long lengths are used ... convenient lengths vary for different types of trail. Each end of flagging is secured to a rock or natural protrusion. Pretty or unusual speleothems are avoided. Flagging is laid in the trail, not on the fragile edges, not on pretties, and not on pristine speleothems. Trails tend to grow out to and sometimes beyond the flagging if given the opportunity. Rocks and protrusions are used as necessary with a wrap and twist of the flagging.

Trails for heavy traffic are laid with about an 18 inch width for walking. This distance allows for easy foot placement when carrying weighty packs; however, more narrow trails work okay to avoid formations on the floor or to encourage slow, deliberate movement through decorated areas. When a stalagmite is obviously used as a handhold, part of it is included inside the flagged trail. Anything inside the flagging eventually is discolored, trampled, or flattened.

Crawl-ways are given two to four feet of trail width, depending on how low the ceiling is and how wide cavers need to spread hands and knees to avoid scraping the ceiling with packs.

On climbs, the flagged trail should be wide enough to accommodate handholds and footholds for cavers with different climbing styles. Cavers move through passages with varying levels of skill and footwork. It helps to try out the trails laid on slopes and climbs, going both up and down, before deciding where to put flagging. The goal is to keep hands and feet inside of flagged routes.

On steep slopes where travel tends to create erosion, zigzagging the path helps preserve the trail and sets the stage for easier travel as the trail becomes worn. On unavoidable steep slopes, safety requires that we remove and set outside the trail any loose rocks that may slide.

Cavers are wearing surgical gloves when flagging trails. Diana Northup has advised that our bare hands or caving gloves may introduce nutrients down lengths of trail that are defining pristine, untouched areas of the cave. On the restoration trips into Lechuguilla, we try to establish practices that are in tune with updated knowledge and that minimize impact on microbial communities. Another consideration is the flagging tape itself. Some flagging tapes are rich in carbon. Are we safe to assume all plastics are inert to cave bacteria? Because some bacteria live on hydrocarbons, more research is needed to better define trail marking materials.

Stray footsteps and trails leading to nowhere are being carefully fluffed. To avoid disturbing surrounding pristine surfaces, each footstep is "fluffed" or lifted only in the impacted area. Small plastic or nylon bristled whisk brooms work well. Occasionally, a fresh plastic spoon is used in unusually delicate areas.

Small laminated paper signs have been used along the trails of Lechuguilla for several years. The Park Service has decided to use additional laminated signs throughout the cave. Part of our restoration task includes listing suggestions for the placement of simple, easy-to-read signs, especially along the trade routes. Examples of signage include: stay inside flagged trails; brush off helmet and clothes before entering this area; boots off here; flowstone shoes only.

RESTORATION TOOLS

On white gypsum surfaces, stainless steel brushes are carefully used with light sweeping motions to peel off layers of grime. A heavy hand will produce unsightly grooves, so special care and a light touch are both pertinent. Stainless steel bristles are necessary because we want to avoid introducing rust and organic materials. Stray bristles that come loose are carried out of the cave. Gypsum debris is caught in plastic bags to avoid contaminating pools or speleothems.

Campsites are fluffed with nylon or plastic bristled whisk brooms before exiting the cave. Care is taken to not stir up dust. By gently brushing the surfaces of the impacted campsite, buried debris and trash is found and carried out. This helps reduce the spreading of campsite edges into pristine surfaces.

Restoration tools that have served the light and packable needs of expedition caving are listed: collapsible or folding buckets with handles; industrial sprayers that come with plastic screens on the end of the sprayer tube, cut off to screw into skinny lightweight plastic water bottles; blue tight-celled car washing sponges designed to stay soft and absorbent (blue pieces are easier to pick up when the sponge begins to deteriorate and they have to be replaced for each trip); hand-sized upholstery brushes found in auto supplies; fresh toothbrushes; whisk brooms with plastic or nylon bristles; powder-free surgical gloves; flagging tape for trails and for special areas; variety of syringes, tweezers, and hard plastic picks; fresh turkey basters; and plastic zip-closure freezer bags, especially the 2 gallon size.

REDUCE CAVER IMPACTS

Continually trying to identify ways to lessen caver impact on Lechuguilla and other caves, we have become aware of some simple actions that can reduce impact. We consciously work at keeping hands off walls, ceilings, and handy leaning places. Obviously, sometimes a handhold is needed for balance; so we try to use small points of contact, knuckles, fingertips, etc., rather than big muddy palm prints. Lots of convenient protrusions and features are along cave trails, but we do not have to use them as rest stations. We can sit within the trails. Why continue to destroy something outside the trail just because somebody sat there first? The old ethic of placing our feet in someone else's footsteps has propagated too much unnecessary impact.

When we see a set of footprints across a cave floor, we try to remove, fluff, or erase the evidence. If we are going to do more damage by trying to restore a footstep, we leave it alone, perhaps place a sign, and hope people will learn that one footprint should not invite more.

Cave softly ... and leave no trace. Move gently. Spot each other through delicate areas. Make it okay to remind each other when we forget and perform some old habit. Think about eliminating the need for restoration. Realize that different types of caves deserve different attitudes. Figure out what the caves in your area need, talk among yourselves, and make it okay to change your behavior.

With respect for the fragile, non-renewable resources of Lechuguilla, we have worked with scientists, cavers, and the National Park Service in composing a set of guidelines for travel in Lechuguilla. Both novice and experienced cavers participate in these trips; thus, some points may seem obvious to the seasoned expedition caver. The following guidelines would not be appropriate for all expedition caves. However, the ideas might provide beneficial protocol for some newly discovered caves around the globe.

MINIMUM IMPACT CAVING IN LECHUGUILLA

The overall goal of Carlsbad Caverns National Park and the Cave Resource Office is to allow limited access to Lechuguilla Cave while minimizing all impacts to the cave. Every person entering the cave is responsible for his/her own actions and for the actions of team members. Expedition Leaders and Team Leaders have tremendous responsibility for the safety and ethics of their personnel and for impacts to the cave. If problems persist, the Leader must abort the trip and the team will leave the cave.

As we learn more about the cave environment, we are in a continuing process of evaluating and redefining caver ethics. The following statement of conduct for Lechuguilla comes from the experiences and thoughtful contributions of many cavers. In any cave, think safety, take care of yourself and your team. Move with stewardship, avoid biological as well as microbial impacts, and take care of the cave resources.

In summary, we all participate in a continuing cycle of learning better ways to protect fragile cave resources. Microbial studies and restoration trips have helped in evaluating the behaviors and ethics that are beneficial in protecting and conserving some types of caves. When developing guidelines for specific caves, or when doing restoration in any cave, it is extremely important to consult with the scientists and managing agencies. In choosing our actions, the cave comes first.

Restoration projects require dedication, time, expense, and a variety of skills contributed from many cavers. Some give volunteer effort. Some give professional committment. All give a heartfelt sensibility for protecting, conserving, and preserving irreplaceable cave resources. The list will continue to grow, but for 1996, on behalf of Lechuguilla and the National Park Service, we thank the following cavers who participated or consulted in the Southwest restoration expeditions. We also thank LEARN cavers for coordinating two weeks of restoration expeditions to the West and the Near East with more than 20 volunteer participants from around the nation. Names include Frank Deckert, Dale Pate, Jason Richards, Harry Burgess, Diana Northup, James Cokendolpher, Rane Curl, Harvey DuChene, Larry Mallory, Jim Werker, Val Hildreth-Werker, Tom Mayer, Joe Miracle, Billy Hoffman, June Hoffman, Barbe Barker, Paula Bauer, Lois Bergthold, Dan Dennison, Mark Esco, Jeff Greving, Carol Holsey, David Kelton, Barry Loucks, Rita Loucks, John Lyles, Bobby Mack, Madina Mansur, Mike Mansur, Andy Mayer, Dan Orr, Roland Vinyard, Dale Webb, Buddy Welker.


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